Usually means: Flies lacking calypteres on wings.
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We found 3 dictionaries that define the word acalyptrate:

General (3 matching dictionaries)
  1. acalyptrate: Merriam-Webster
  2. acalyptrate: Wordnik
  3. acalyptrate: Wiktionary

Definitions from Wiktionary (acalyptrate)

adjective:  Of or pertaining to the Acalyptratae group.
noun:  A muscoid fly belonging to the subsection Acalyptratae.

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